Permanent change in the structure of root ,to carry new function,suitable to the changed environment is called Root modification.Entire root or a part of it may be modified.Various Root modifications are:
1. Tuberous or storage roots: Due to storage of food ,roots become bulged.,changed in shape. Due to storage tap roots are spindle shaped in Raphanus sativus,Conical in Daucus carota,Napiform in Beta vulgaris. These roots store nutritive materials and are called root crops. These plants are called Biennials growing for two years or two seasons. Plants prepare and store food in the tap roots at the end of first year and utilize the food in the second year producing flowers and fruits. Root crops should be harvested at the end of first year only. In Ipomoea batat a single tuberous root,in Asperagus a cluster of tuberous roots are found. In Beta vulgaris food is in the form of sugar, in Dahlia in the form of Inulin, in all others in the form of starch.
2. Epiphytic (or) Velamen Roots: The plants taking shelter on the branches of other tall trees for more sunlight are called Epiphytes. They have two types of roots,a)branched short clinging roots fix epiphyte on the branch of host plant.
long brown branched freely hanging Velamen roots absorb moisture from atmosphere.These roots contain dead hygroscopic epidermal tissue called Velemen.Epiphytes show xerophytic features as they do not get continuous water supply. Ex: Vanda.
3. Photosynthetic (or) assimilatory roots: In epiphyte Taeniophyllum,stem,leaves are absent.Roots are aerial,flattened,green,synthesizing food and are called assimilatory roots.In Tinospora,aerial roots,in Trapasubmerged roots are green with chloroplasts.
4. Pneumatophores (or) respiratory roots: Mangroves growing in saline,marshy,water logged soils do not get oxygen for root respiration from soil. Some special aerial branches from roots form respiratory roots Pneumatophores with small pores Pneumathodes for gaseous exchange.ex:Rhizophora, Avicennia.In free floating hydrophytes Jussia,breathing roots are spongy,
5. Parasitic roots(or) haustoria: Plants depending on other plants completely or incompletely are called Parasites.Special adventitious rootsproduced by parasites to get food and nutrientsfrom host plant are called Haustoria.
Parasites are a)stem parasites(on host stem) are complete parasites ex:Cuscuta,and in- complete parasites ex:Viscum.b) root parasites (on host root ) are complete root parasites ex:Orobanche and in-complete root parasite ex:Striga.Leaf less ,nonchlorophyllous parasites getfood and mineral water from the host plant by establishing haustoria with both phloem and xylemforming complete parasites.In-complete parasites have green leaves,establishing haustoria with xylem of host plant to get mineral water only.
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AERIAL STEM MODIFICATIONS.
When the entire or a part of the aerial stem changes into a permanet structure to carry another function in changed environment it is called aerial stem modification.
1. Tendrils: Tendril is delicate wiry coiled sensitive organ useful for climbing.Tendril coils around the supporton contact.In Passiflora axillary buds, in Vitis terminal buds develop into tendrils
2. Thorns: Hard woody pointed vascularised structure meant for protection is called thorn.In Bougainvilla axillary bud,in Carissa terminal bud (into a pair of thorns) modifies.In Duranta axillary bud develops into thorn bearing leaves and flowers.In Punica granatum axillary bud develops into thorn bearing leaves and branches.
3. Hooks: Sensitive curved woody structure helping in climbing is called hook.Axillary bud forms hook inHugonia.In Artabotrys terminal bud forms inflorescence whose peduncle curves forming hook.
4. Phylloclade: Leaf-like stem is called Phylloclade.It is a xerophytic feature.To reduce transpiration leaves modify into spines,stem becomes green with chlorophyll forming Phylloclade.In Opuntia leaves of axillary bud form a group of spines some may be small hair like barbs. Group of spines and barbs at each node represent areole.Flattened short green or cylindrical branches of limited growth are called Cladophylls or Cladodes.They are cylindrical ,one internode length formed from axils of leaves in Asparagus.In Ruscus they are flat leaflike formed from axils of scale leaf. Chladophyll bears male or female flowers at node on the surface.
5. Tuberous stems: Aerial stem becomes tuberous due to storage of food material.In Bulbophyllum an epiphyte, one basal internode forms Pseudobulb due to storage of food.In Brassica oleracea var gangyloides (knoll-khol) entire stem becomes tuberous due to storage of food.
6. Bulbils: The modified condensed branches with stored food helping in vegetative propagation are called Bulbils.In Dioscorea bulbifera bulbils form from the axils of bracts.In Oxalis a group mof bulbils develop from the top of tuberous root.
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SUB-AERIAL STEM MODIFICATIONS.
These are weak stemmed plants. Stems are partly aerial and partly underground.
These are 4 types.
1. Runners; ex:Oxalis; Lippia; Stem creeps on the soil rooted at every node. Nodes can grow into independent plants (vegetative propagation),when the internodes break off.
2. Stolons;ex:Nerium,Jasminum. These are basal long branches growing obliquely ,producing adventitious roots when touch the soil.These stolons can become independent plants when separated from the main plant.(layering process)
3. Suckers: ex:Mentha;Chrysanthemum. These are obliquely growing underground
Branches of the stem.These suckers with adventitious roots below.can become independent,when separated.
4. Offsets: ex
istia;Eichhornia.In these free floating hydrophytes,stem is very much reduced,leaves form in rosette.Axillary buds develop into short slender branches of one internode length called Offsets.From the tip of the offset leaves form above,roots below.Offsets become independent plants when they are cut from the main plant.Agave Americana terrestrial land plant produces offsets.
UNDERGROUND STEM MODIFICATIONS
These are stems growing inside the soil.They are fleshy swollen,due to food storage.They are protected from herbivorous animals.They help in vegetative propagation and perennation.So these are also called multipurpose stem modifications.These are:
1. Rhizome:ex:Zingiber officinalis,Canna indica. Horizontally growing underground stem is called Rhizome.It has nodes,internodes,scale leaves at nodes,adventitious roots below.shoots above.Rhizome perennates even in drought.
2. Corm: ex:Colacasia; Amorphophalus. Vertically growing underground stem at particular depth is called Corm.It is tuberous with nodes,internodes,scale leaves.
Axillary buds form daughter corms.Apical bud forms aerial shoot.The special adventitious roots keeping the corm at particular depth in Amorphophalus are called Pull roots or contractile roots.
3. Stem tuber: Ex;Solanum tuberosum.The tips of the underground branches become
tuberous due to storage of food are called Stem tubers.It has outer brown coloured
periderm,eyes.Each eye-like structure has semilunar leaf scar with axillary bud
representing node.It helps in vegetative propagation.Stem tuber has no roots.The
stored food is starch in Solanum tuberosum, inulin in Helianthus tuberosus, Stachyose
in Stachys tubifera.
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4. Bulb: Special type.Stem is reduced to biconvex disc,with roots belowleaves above the soil.Leaf bases become fleshy with food and water.Terminal bud grows into aerial
inflorscence,axillary buds produce daughter bulbs,helping in vegetative propagation.
Bulb is only the underground stem that does not store food in stem.Based on the
arrangement of scale leaves bulbs are two types.
a) Tunicated bulb: ex:Allium cepa.Fleshy leaf bases overlap in concentric rings,circles.Entire bulb is covered by dry membrane called tunic.
b) Scaly bulb: Allium sativum.Imbricate or naked bulb.Individual scale leaves become fleshy due to storage of food.They are loosely arranged as in Lilium candidum and fleshy cloves are enclosed by whitish skinny dry tunic as in Allium sativum.
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LEAF MODIFICATIONS
If the entire leaf or a part of it undergoes permanent change to perform new
function suitable for the environment it is called leaf modification.They are:
1. Tendrils: Weak stemed plants climb up the support by delicate wiry tendrils.In Lathyrus entire leaf except stipules,in Pisum terminal leaf lets of compound leaf,in Smilax stipules, in Nepenthes part of petiole, in Clematis petiole modify into tendril.
2. Spines: Sharp pointed structures.Xerrophytic character.Spines reduce the rate of transpiration.They protect the plants from herbivorous animals.In Asparagus entire leaf, in Phoenix leaf tip, in Acacia stipules , in Argemone leaf margin,in Citrus first leaf of axillary bud modify into spines.
3. Scale leaves:Thin small colourless,dry membranous,structures are called scales.In Casuarina phylloclade scales reduce transpiration.In Zingiber underground stem scale leaves protect axillary and terminal buds . In Allium scaleleaves store food.
4. Phyllode:green flattened photosynthetic petiole or secondary rachis is called Phyllode.In Acacia melanoxylon,petiole of the pinnately compound leaf modifies into phyllode.In Parkinsonia primary rachis ,stipulesmodify into spines.Secondary rachis forms phyllode,with small deciduous leaflets.
5. Reproductive leaves:Leaves with epiphyllous buds are called reproductive leaves.They help in vegetative propagation.In Bryophyllum epiphyllous buds lie in the notches of crenate leaf margin.In Scilla indica leaf tip bears bud.,that grows into new plant when the leaf tip touches the soil.In Begonia injured parts of leaf produce buds.
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6. Trap leaves,insectivorous leaves,carnivorous leaves:These plants grow in Nitrogen deficit soils .Leaves are used as traps to catch and kill the insects to get nitrogen in the form of proteins.Insect proteins are digested and assimilated into the
plant .Insectivorous plants secrete digestive juices in leaves.ex: Drosera, Dionaea,
Utricularia, Nepenthes.
Mechanism in Nepenthes: It is called Pitcher plant.Lower part of petiole is wing like, upper part forms tendril. Leaf lamina forms pitcher .Leaf tip forms lid.Lid is immovable colourful,attractive.Rim of Pitcher is slippary ,lined with digestive
glands and downwardly projected hairs into pitcher.Insect is attracted by lid and nectar.When the insect lands on rim it slips down and drowned in acidic fluid in pitcher.Downwardly projected hairs prevent the insect escape .Insect is killed and digestedby proteolytic enzymes.Nitrogenous substances are absorbed and transported.
RACEMOSE TYPE INFLORESCENCE
Peduncle grows indefinitely. Flowers are arranged in acropetal manner ie.older flowers are at bottom younger ones are at the top. When the peduncle is reduced opening of flowers is centripetal ie. From periphery to center. Two types:
A) Racemose type with pedicellate flowers:
1. Simple Raceme: ex: Crotalaria. Bracteate pedicellate,flowers are arranged in acropetal manner,on indefinitely growing peduncle.
Compound Raceme: Mangifera, Yucca. It is also called Panicle.Peduncle is branched. Each branch resembles a simple raceme.
2. Simple Corymb;ex: Cassia; Gynandropsis.It is like a simple raceme.
But all the flowers are brought to the same level,height. Lower flowers have long pedicels and upper ones have shorter pedicels.
Compound Corymb; ex: Pyrus malus; Brassica oleratia var. botrytis.
Peduncle is branched .Each branch is a simple corymb.In Cauliflower peduncle is fleshy storing food.
3. Simple Umbel : ex: Allium cepa.Many pedicellate bracteate flowers
arise as a cluster at the apex of unbranched condenced peduncle.All the
bracts unite to form whorl called “ involucre”. Flowers open in
centrifugal manner.
Compound Umbel : ex: Coriandrum sativum., Daucus carota. Peduncle
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is branched. Each branch bears a simple umbel.All the branches arise in umbel manner .Involucre of bracts occur at the base of branches.Involucel of bracteoles occur at the base of simple umbels.
Racemose type with sessile flowers:
1. Simple Spike: ex:Achrynthes, Amaranthus. Bracteate sessile flowers arise acropetally on unbranched peduncle.Similar to simple raceme but for the sessile flowers.
Compound spike .( spike-let) ex: Oryza; Triticum. Peduncle is branched.It grows indefinitely.Each branch produces bracteate sessile flowers acropetally.Each branch is a spike-let. Its axis is called Rachilla.Bracts are glumes.Basal glumes without flowers are sterile .Fertile glumes are with flowers.Bracts form lemma, bracteoles form palea,the reduced perianth form lodicules. Characteristic in family
oaceae.
2. Catkin : ex : Casurina ; Acalypha. It is spike like.Peduncle is
long.weak,drooping. Many sessile,unisexual flowers arise on peduncle
in acropetal manner. Often called “ Amentum “ as is found in
Amentiferae.
3. Simple Spadix ;ex: Colocasia. Peducle is fleshy unbranched.The terminal flowerless part is called Appendix.Sessile unisexual flowers are below the appendix,female flowers below and male flowers above, neutral flowers in between.All the bracts unite to form thick leathery Spathe,covering the entire inflorescence.Common in Family.Aroideae.
Compound spadix: ex:Cocos nucifera. Peduncle is branched . On each branch arise unisexual female flowers below,male flowers above in acropetal manner.In Cocos spathe is hard woody boatlike,.In Musa each branch is covered by leathery spathe.
4. Head (or ) Capitulum ( or ) Anthodium : Highly evolved in Racemose
type.Seen in Asteraceae.Peduncle forms a flattened disc called
receptacle.Small sessile flowers are centripetally arranged on this.
Bracts form involucre around the inflorescence.Flowers are two types:
A) Disc flowers: actinomorphic,bisexual,central in position.
Ray
flowers : zygomorphic,unisexual female,periphery in position.Based on the types of flowers present ,head may be Homogamous with only one kind of flowers.ex
nly disc flowers in Vernonia,only ray flowers in Tagetus.Heterogamous Head has both Ray flowers in margin and Disc flowers at center,ex: Tridax,Helianthus
Compound Head :Ex: Echinops; Sphaeranthus. Axis is branched.Each branch ends in Head inflorescence.
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CYMOSE (OR ) DEFINITE INFLORESCENCE.
Flower develops at the ip of the peduncle.Its growth is definite.From the axils of bracts below this terminal flower,branches develop,which also end in flowers.Flowers arrangement is basipetal.ie first formed older flower is at apex,the later formed younger flowers at below.On the condenced axis opening of flowers is centrifugal ie. From center to periphery.
Cymose types are:
1. Solitary Cyme: Simplest type.Axis ends in a flower.Flower is axillary in Hibiscus,terminal in Datura.The articulation between the peduncle and pediceldistinguishes the nature.
2. Simple Cyme: A three flowered inflorescence.Axis ends in a flower.Below this two lateral flowers develop,in basipatal manner.ex:Jasminum, Bougainvillea.
3. Monochasial cyme: Only one branch develops below,when axis ends in a flower.The same nature repeats.Single branches unite forming Sympodial axis,looking like Racemose.In this flowers are not axillary. They are bracts opposed.They are two types on the nature of branches arranged.a) Helicoid:ex:Hamelia. Succesive lateral branches develop on one side,in spiral manner.
Scorpioid:ex:Solanum.successive lateral branchesgrow on either side,in zig-zag manner.
4. Dichasial cyme:ex: Ipomoea.Two lateral branches develop below ,when axis ends in terminal flower. Inturn each lateral branch ends in a flower repeating the same process. This is called Dichasial as two branches develop at each time.(Clerodendron).
5. Polychasial Cyme: ex: Nerium.Axis ends in flower . Many lateral branches develop below.Each ends in flower.This nature continues.It is called Polychasial,since more than two lateral branches develop.
SIMPLE FRUITS.
Fruit developing from syncarpous gynoecium of a single flower is called simple fruit.On the nature of pericarp,simple fruits may be :
FLESHY FRUITS: Pericarp is fleshy,at the time of ripening.Separable into outer epicarp,middle mesocarp and inner endocarp.Seeds are liberated out when pericarp decays.
1. Berry: Fruit develops from bicarpellary or multicarpellary syncarpous gynoecium.Pericarp is separated into outer thin epicarp;middle mesocarp and inner endocarp fuse to form pulp with numerous seeds.ex: Solanum melongena.In Musa
berry is developed from inferior ovary.In Phoenix dactilifera single seeded berry
develops.
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2. Pome:ex: Pyrus malus.Fruit is developed from bicarpellary or multicarpellary gynoecium,inferior ovary. Fleshy thalamus around the gynoecium is fleshy.Pericarp is hard enclosing the seeds.
3. Pepo:ex: Cucumis. Fruit is developed from uni or tricarpellary syncarpous gynoecium,inferior ovary.Pericarp is divided into outer epicarp fused with hard thalamus forming rind,fleshy mesocarp,smooth endocarp.Seeds are developed from the inner walls of pericarp.
4. Hesperidium:ex:Citrus species. (Rutaceae) Fruit is developed from multicarpellary syncarpous multilocular superior ovary. Pericarp is divided into outer leathery glandular epicarp,white papery mesocarp,many chambered inner endocarp with juicy edible hairs.Seeds on central axis.
5. Drupe:ex Cocos nucifera;Mangifera indica. Fruit develops from uni or multicarpellary superior ovary,unilocular,one seeded,with stony endocarp.In Mangifera,epicarp is leathery,mesocarp is fleshy,juicy,edible,endocarp is stony.In Cocos,epicarp is leathery,mesocarp is fibrous,endocarp is hard stony.Endosperm of seed (copra)forms edible part.
DRY FRUITS.: Pericarp is dry, undifferenciated when ripe.
DRY DEHISCENT FRUITS.
Pericarp breakes liberating seeds at maturity.They are:
1. Follicle:ex:Calotropis.Develops from superior ovary.Dehisces along one suture only.
2. Legume:ex
olichos.Develops from monocarpellary unilocular half superior ovary.Fruit breaks dorsiventrally into two halves liberating the seeds.
3. Siliqua:ex:Brassica.Developed from bicarpellary syncarpous unilocular ovary but made bilocular by false septum called replum.Seeds are on either side of replum.Pericarp breakes dorsiventrally from base to apex leaving seeds on replum.
4. Silicula;Ex:Capsella bursa pastoris.Short broad siliqua with limited number of seeds.
5. Capsule: Developed from uni or multicarpellary mulilocular superior ovary.
.Based on dehiscence, capsule may be:
a) Septicidal capsule:ex:Aristolochia.Capsule breaks along the septae,in between
locules,leaving seeds within the valves
b) Loculicidal Capsule:ex : Abelmoschus esculentus.Capsule dehisces through the middle portions of locules.Number of pieces are equal to the number of carpels.
c) Septifragal Capsule:ex: Datura. Pericarp dehisces either by loculicidal or septicidal leaving seeds on swollen axis of placenta,at the center.
d) Porous Capsule:ex: Papaver.Fruit is cup like with lid.Many pores in between on margin.Seeds are dispersed through the pores present on upperside..In Campanula from inferior ovary with pores at base.
e) Pyxidium:ex
ortulaca.Spherical shaped capsule developed from superior ovary.Pericarp splits transversly.Upper lid falls off leaving seeds on basal cup like structure.
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DRY INDEHISCENT FRUITS
One seeded. Never break even at maturity.Seed is liberated only after the decay of the pericarp.These fruits are:
1. Achene: ex:Clematis. Developed from monocarpellary unilocular superior ovary.Pericarp and seed coat are well separated.
2. Caryopsis:ex: Oryza. Pericarp and seed coat are fused together.
3. Cypsela: ex: Tridax. Developed from bicarpellary syncarpous unilocular inferior ovary.Persistant calyx form pappus helping in fruit dispersal.
4. Nut: ex: Anacardium.Developed from Bi or multicarpellary syncarpous unilocular superior ovary.Pericarp and seed coat are free,separated.Pericarp is hard,stony.
5. Samara: ex: Winged fruits.Developed from bi or tricarpellary syncarpous superior ovary.Wings develop from pericarp in Hiptage,from caly in Gyrocarpus,from style in Ventilago,from calyx and style in Shorea.
SCHIZOCARPIC FRUITS.
Fruit breaks into many single seeded bits called mericarps.Fruit shows both dehiscent and indehiscent features like breakage of pericarp,degeneration of pericarp to liberate seeds.Various types are:
1. Lomentum: ex: Acacia; Mimosa. Developed from monocarpellary gynoecium. Fruit is constricted inbetween the seeds,into which fruit breaks at maturity.Common in Mimosae.
2. Schizocarp:ex: Abutilon; Sida.Developed from multicarpellary syncarpous multilocular superior ovary.Fruit breaks into many mericarps.Each mericarp may have one seed ( Sida) or many seeds ( Abutilon ).Common in Malavaceae.
3. Carcerulus:ex: Ocimum. Developed from bicarpellary syncarpous tetralocular ovary.Fruit breaka into four nutlets.Each mericarp has stony pericarp.
4. Regma:ex: Ricinus. Developed from tricarpellery syncarpoustrilocular superior ovary. At maturity fruit breaks into three cocci,one seeded mericarps.
5. Cremocarp:ex: Coriandrum. Developed from bicarpellary syncarpous bilocular inferior ovary.At maturity fruit breaks into two one seeded maricarps attached to carpophore,an elongated thalamus.Common in Apiaceae( Umbelliferae ).
6. Double Samara:ex:Acer. Developed from bicarpellary syncarpous superior ovary.Fruit breaks into two samara like mericarps.
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TAXONOMY
MALVACEAE
Examples: Systematic Position:
Abelmoschus esculentus ( benda) Class: Dicotyledonae.
Gossypium herbaceum (patti) Sub.Class: Polypetalae.
Hibiscus rosasinensis. (mandara ) Series: Thalamiflorae.
Hibiscus cannabinus (gongura ) Order: Malvales.
Thespesia populnea ( gangaravi ) Family: Malvaceae.
Vegetative Characters:mesophytes,mostlt herbs or shrubs,a few are trees (Thespesia ),
Vegetative parts are covered by stellate hairs,plant tissue contain mucilage cavities,mostly simple leaf (Hibiscus),palmately lobed ( Gossypium ),reticulate venation.
Floral characters:
Inflorescence:Solitary cyme,axillary in Hibiscus,terminal in Gossypium
Flower: large showy bracteate,bracteolate,3-10 bracteoles form a whorl called
Epicalyxoutsidecalyx,epicalyxisabsentinSida,Abutilon,bisexual,pentamerous,hypogynousCalyx: 5,gamosepalous,valvate.
Corolla: 5,polypetalous,twisted,fuse at base with staminal tube.
Androecium: numerous,epipetalous,monadelphous in Hibiscus,form staminal tube around style,anthers monothecous,reniform,extrorse,trnsverse dehiscence,large apherical pollen,with spinous exine.
Gynoecium: 5,in Hibiscus,syncarpous,ovary superior,number of locules equal to number of carpels,ovules on axile placentation,simple terminal style,stigma lobes equal to number of carpels in Hibiscus.
Floral Formula: Br,Epik3-10,+,o+’,K(5),C5,~A(&),G(3-&)
Pollination: cross pollination is due to protandry,flowers entamophilous.
Fruit: loculicidal capsule in cotton,schizocarp in Abutilon,berry in Malvaviscus.
Seed: nonendospermic,seedcoat has hairy outgrowths (cotton).
Economic importance:
1. fibre yielding,leafy vegetable-Hibiscua cannabinus.
2. Vegetable-Abelmoschus esculentus
3.cotton sps.-cotton yielding,cotton seed oil in soap making,oil cake as
cattle feed.
4. medicinal plants-Abutilon,Sida.
Important characters:1)stellate hairs on vegetative plant parts,with mucilage within tissues,2)epicalyx, 3)stamens=numerous monadelphous,staminal tube,anthers monothecous,reniform.4) superior ovary,number of carpels equal to number of locules in ovary,axile placentation, 5) fruit=capsule.
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FABACEAE.(sub.fam. Papilionaceae of fam.Leguminaceae.)
Examples: Sys.Position:Cla: Dicotyledonae.
Arachis hypogea (verusenaga ) Sub.Clas
olypetalae.
Cajanus cajan (kandulu ) Series: Calyciflorae.
Cicer arietinum (sanaga) Ord: Rosales.
Dolichos lab lab (chikkudu ) Fam: Fabaceae.
Glycine max (soya bean )
Phaseolus mungo (minumulu ),Phaseolus aureus (pesalu),Pisum sativum (batani)
Veg. Characters: mesophytes,plants show great diversity in habit,twainer (Dolichos) tendrillar climber (Pisum),root nodules with Nitrogen fixing bacterium Rhizobium with pink pigment leg-haemoglobin.
Leaf: cauline,alternate,stipulate,foliacious in Pisum,pulvinate leaf base,simple.pinnately,trifoliate compound leaf(Dolichos),entire leaf in Lathyrus,terminal leaf lets in Pisum modify into tendrils,leaflets into spines in Ulex.
Floral characters:inflorescence axillary or terminal,raceme (Crotalaria ),paniculate.
Flower: bracteate.bracteolate,pedicellate,complete,bisexual,pentamerous,perigynus,thalamus cup shaped,zygomorphic due to Papilionaceous corolla.
Calyx: 5,gamosepalous,valvate,odd sepal anterior in position.
Corolla: 5, free, Papilionaceous with bigger posterior Vexillum,two lateral Alae,two boat shaped Carina towards anterior side enclosing the stamens ,gynoecium,decendingly imbricate.
Androecium: 10,didelphous,9+1 in Dolichos,monadelphous in Crotalaria,anthers dithecous,introrse,longitudinal dehiscence.
Gynoecium: monocarpellary,unilocular,half-inferior,superior(Crotalaria),many ovules in two vertical rows on marginal placentation,long style, simple stigma,nectar gland at base.
Floral Formula:Br,Brl,%,O,K(5),C1+2+(2),A(9)+1,G1-
Pollination: Protandrous favouring croos pollination.Piston mechanism-essential organs move like Piston,when the insect alights on wings,pressed them,keels open, stigma comes out receiving pollen,then filaments come out shedding pollen on insect.
Fruit:legume in Dolichos,geocarpic in Arachis,samara in Pterocarpus,Dalbergia.
Seed: nonendospermic,dicotyledonous,store proteins in large amounts.
Economic importance:1)pulses are rich source of proteins.2) pods are vegetables
Ex
olichos, Glycene.3)Ground nut oil from Arachis hypogea,Soya bean oil from Glycene max.used in cooking 4)fodder from Crotalaria, Phaseolus.5)fibre(sunhemp) from Crotalaria juncea.6) seeds as condiment,leaves as vegetable ex:Trigonella.7)blue dye,Indigo from Indigofera tinctoria 8) musical instruments from wood of Pterocarpus santalinus.9)furniture from wood of Dalbergia latifolia.
Imp. Characters: 1) roots –nodular,help in N2 fixation.2)leaves-stipulate,pulvinate,show sleeping movements,3)flower-perigynus,thalamus cup shaped,odd sepal anterior,Papilionaceous corolla,descendingly imbricate aestivation,odd petal posterior,androecium 10 didelphous,Gynoecium monocarpellary,marginal placentation,4)Fruit- legume or pod.
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ASTERACEAE: most evolved family in Dicots,Being Aster as nomenclatural type,with suffix acae(absent in Compositae),named as Asteraceae as per ICBN.
Examples: Sys.Position:Cla: Dicotyledonae.
Carthamus tinctorius(safflower) Sub.Cla: Gamopetalae,
Chrysanthemum indica(chamanti) Series: Inferae,
Cichorium intybus(chicory) Ord: Asterales,
Eclipta alba(bringaraj),Tagetus erecta(banthi) Fam: Asteraceae.
Helianthus annuus (sunfliwer),Tridax procumbens(gaddi chamanti)
Veg. Characters: fasciculated tuberous roots in Dahlia and Cichorium,herbaceous stem contains oil passages and latex,prostrate decumbent(Tridax),Launea is runner,in Helianthus tuberosus undergroun tuber stores inulin,leaves simple covered by hairs,compound in Dahlia.
Floral characters: head inflorescence highly evolved,homogamous with Disc florets only in Vernonia,with Ray florets only in Chrysanthemum,heterogamous with central Disc florets surrounded by peripheral Ray florets in Tridax.
Disc floret: central sessile,bracteate,actinomorphic,complete,pentamerous,epigynous.
Ray floret: peripheral,sessile,bracteate,zygomorphic due to ligulate corolla,incomplete,unisexual female.pentamerous, epigynous.
Calyx: reduced to pappus,persistent,absent in Xanthium,helps in fruit dispersal.
Corolla
isc=5,gamopetalous,tubular,valvate. Ray=5,two reduced ,three form tongue shaped ,gamopetalous,valvate,nectar glands at base.
Androecium: Disc= 5,epipetalous,syngenesious(filaments free,anthers united) hooded,dithecous,introrse,longitudinal dehiscence. absent in Ray floret.
Gynoecium:similar in both Disc &Rayflorets, bicarpellary, syncarpous, unilocular, inferior,single ovule on basal placentation.style terminal,stigma bifid,passes through united anthers.
Floral formula: Disc floret:Br,Ebrl,+,O,K pappus,C(5),A(5),G(2)_
Ray floret:Br,Ebrl,%,O,Kpappus,C(5),A0,G(2)_
Pollination: protandrous,entamophily,anemophily in Xanthium,safty mechanism(bending of stigma lobes receiving self pollen, when cross pollination fails).
Fruit: single seeded cypsela,persistent pappus provide parachute mechanism. In Xanthium hhoklike spines help in dispersal by animals.
Economic importance:1) stem tubers of Helianthus tuberosus,root tubers of Dahlia are edible.2) Lactuca sativa is leafy vegetable.3) from seeds of Carthamus tinctorius safflower oil,Helianthus annues sunflower oil is extracted. 4) chicory from the roots of Cichorium intybus.5) antihelmenthic drug Santanine from the flower heads of Artemisia vulgaris. 6) Eclipta alba used in hair tonic.7) insecticide Pyrethrum from flowers of Chrysanthemun cinerariaefolium.
Imp.Characters(advanced characters):1) plants-herbaceous,veg.parts with internal oil cavities.2)head-homogamous,heterogamous,reduced florets,3) calyx
appus persistent,helps in fruit dispersal,4) stamens: epipetalous,syngenesious,hooded,
5) gynoecium
icarpellary syncarpous,unilocular,inferior ovary,single basal ovule.6)fruit:Cypsela,parachute mechanism.
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SOLANACEAE.
Examples: Sys.Position:
Atropa belladonna (belladonna) Class: Dicotyledonae
Capsicum fruitiscens (chilly) Sub.cla: Gamopetalae
Cestrum nocturnum (night queen ) Series: Bicarpellatae
Cestrum diurnum ( day king ), Order
olemoniales,
Datura metal (thorn apple) Fam: Solanaceae.
Lycopersicon esculentum (tomato ) Nicotiana tabacum (tobacco)
Solanum melongena (brinjal ), Solanum tuberosum (potato ), Withania somnifera (aswagandha )
Vegetative characters:Solanum surattense is xerophyte,stem tuber in Solanum tuberosum,adnation of petiole with stem,Internally stem shows bicollateral vascular bundles(phloem on either side of central xylem,separated by cambia),petiole adnates with internode,simple pinnately lobed leaves.
Floral characters:
Inflorescence: cymose,axillary,terminal in Datue,extra axillary scorpiod cyme.
Flower: bracteate,ebracteolate,bisexual,pentamerous,hypogynous,obliquely zygomorphic due to rotate carpels.
Calyx: 5,gamo,valvate,persistent,macrescent(does not grow after fertilization),ex
rinjal, accrescent(grow along with fertilized ovary after fertilization)ex
hysalis.
Corolla:5,gamo,valvate or plicate.
Androecium: 5,epipetalous,anther large,dithecous,longitudinal dehiscence in Datura, porous dehiscence in Solanum.
Gynoecium: bicarpellary syncarpous bilocular, made tetralocular by false septum,ovules on axile placentation,ovary oblique,posterior carpel to right side,anterior carpel to left sideat 45 anglemade the flower obliquely zygomorphic,terminal style,capitate stigma.
Floral Formula: Br,Ebrl,+,O,K(5),C(5),A5,G(2)_
Pollination
rtandrous,protogynous(Solanum),entemophily,self pollination (Nicotiana).
Fruit: berry in brinjal,tomato.chilly,septifragal capsule in Datura,Nicotiana,
Eco.Imp: 1) Vegetables-potato(Solanum tuberosum),brinjal(Solanum melongena),
Tomato(Lycopersicum esculentum),chilly(Capsicum fruitiscens)2)alkaloid-capsine
From Capsicum used as condiment.3) alkaloid-Nicotine-from leaves of Nicotiana. 4) atropine-from Atropa belladonna used in plasters.5) Solanum surattense,Datura stramonium,cures asthma,6) Withania somnifera-root extract used as rejuvenating tonic.
Imp. Characters: 1) bicollateral vascular bundles in stem,2) adnation of petioles with axis.
3)persistent calyx.4) bicarpellary syncarpous,bilocular made tetralocular,by false septum,ovules on swollen axile placentation,5) obliquely placed superior ovary,due to rotation,6)fruit berry or capsule.
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LILIACEAE:
Examples: Sys.Position:
Allium cepa ( onion ) Class: Monocotyledonae
Allium sativum (garlic ) Series: Coronariae,
Aloe barbadensis (aloe ) Family : Liliaceae.
Asperagus racemosus (sathamuli ),Colchicum autumnale
Ruscus aculeate,Smilax zeylanica,Yucca gloriosa.
Vegetative Characters: mesophytes (Allium ),xerophytes ( Ruscus ),perennial shrub (Aloe ),climbers ( smilax ),fasciculated tuberous roots (asparagus ),bulb (Allium) Rhizome ( Gloriosa ),corm ( Colchicum ),naked bulb (Lilium ),tendrillar climber (Gloriosa, Smilax ),In Yucca, Dracaena aerial stem shows anamolous secondary growth,cladophylls ( Ruscus,Asparagus ),vegetative propagation by axillary buds ( Lilium bulbiferum ),suckers ( Aloe ).
Leaves: radical (Allium),cauline (Smilax),parallel venation, reticulate ( Smilax) succulent leaves (Aloe, Yucca),scales ( Asparagus,Ruscus),leafbases fleshy (Allium,Lilium ),In Gloriosa leaf tip, in Smilax stipules modify into tendrils,
Leaf apex with epiphyllous buds in Scilla.
Floral Characters:
Inflorescence:racemose,simple raceme in Asparagus,panicle in Yucca,umbel in Allium,solitary terminal in Lilium,axillary in Gloriosa,stalk like peduncle emerging from underground stem is called scape.ex: Allium.
Flower: bracteate,ebracteolate,pedicellate,actinomorphic,trimerous,bisexual,hypogynous, unisexual in Ruscus,Smilax.
Perianth:6 tepals,3+3 in two whorls,polyphillous in Lilium,gamophyllous in Aloe,petaloid,odd petal is anterior in position,valvate.
Androecium: 6,3+3,free,epiphyllous,dithecous,introrse,longitudinal dehiscence.
Gynoecium: tricarpellary syncarpous trilocular,ovules on axile placentation,superior ovary,terminal style,trifid stigma,capitate.
Floral formula: Br,Ebrl,+,O,P3+3,A3+3,G(3)_
Pollination:entamophilous,protandrous in Allium,protogynous in Colchicum,Pronuba yuccasella, moth associates in Yucca,herkogamy in Gloriosa.
Fruit: berry in Asparagus,loculicidal capsule in Lilium,septicidal capsule in Gloriosa.
Seed: monocotyledonous,endospermic,polyembyony in Allium.
Eco.Imp: 1) bulbs of Allium edible,bactericidal 2) cloves of Allium sativum spice,medicinal curing gastric,heart complaints,3) fasciculated tuberous roots of Asparagus are edible.4) Aloe treats piles.5) roots of Smilax source of Sarsaparilla medicine,6) colchicines,chemical mutagen,from corms of Colchicum autumnale.
Imp.Cha: 1) perennial underground stems with fibrous roots 2) simple leaves parallel venation3) flowers actinomorphic,trimerous,perianth homochlamydeous,petaloid,4)Gynoecium-tricarpellary syncarpous,trilocular,ovules on axile placentation,superior ovary,5)fruit capsule.
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INTERNAL ORGANISATION
INTERNAL STRUCTURE OF DICOT STEM. Ex: Helianthus annuus( sunflower )
T.S of primary Dicot stem shows three zones:
I) epidermis: Outer most layer. Rectangular cells are arranged compactly without inter cellular spaces.Cuticle deposited on the outer surface is made up of waxy substance cutin.Stomata are peresent. Multicellular uni or biseriate trichomes develop on epidermis. Cuticle, trichomes check water evaporation.They protect the stem from high temperature.Trichomes prevent the entry of pathogenic organisms Through stomata ,epidermis allows exchange of gases and promotes transpiration.
II) Cortex: It is the part between epidermis and stele. It shows:
1) Hypodermis: Outermost part of cortex,beneath epidermis.,made up of 3 to 6 rows of collenchyma.The cells are compoactly arranged without intercellular spaces,due to excessive thickening of cell corners.It gives elasticity to the stem. By possessing chloroplasts it helps in the production of food material.
2) General cortex: Below the hypodermis,5 to 10 rows of parenchyma are present.The cells are thin walled isodiametric,with or without intercellular spaces.Outer layersd contain chloroplasts,inner layers with leucoplasts,performing assimilation and storage respectively.
3) Endodermis:Innermost layer of cortex.Cells are barrel shaped,compactly arranged without intercellular spaces.Radial and transverse walls show lense shaped thickenings called casparian bands.Endodermis is called starch sheath as it contains starch grains.
III) Stele:central conducting cylinder. It is well developed. It consists of :
1) Pericycle: It is between endodermis and vascular bundles.It is formed of Sclerenchyma,in the form of semilunar patches,intervening with parenchyma masses.
2) Vascular bundles: 15 to 20 in circular ring.The arrangement of vascular bundles in a ring is called Eustele.It is concerened with conduction of water,salts and food materials.Each vascular bundle is top shaped ,phloem outerside,xylem innerside,with meristem inbetween(fascicular cambium) on the same radius. Vascular bundle is called conjoint (xylem and phloem form one bundle ),collateral (xylem and phloem on same radius),open(cambium inbetween xylem and phloem ). Xylem is endach (protoxylem towards center).Xylem has vessels,xylem parenchyma,xylem fibres.Phloem has sieve tubes,companion cells,phloem parenchyma.
3) Medulla: central part of stele.It is filled with parenchyma storing food material.
4) Medullary rays:Thin walled living,radially elongatedfrom medulla to periphery,in between the vascular bundles are called medullary rays.They help in lateral conduction
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INTERNAL STRUCTURE OF MONOCOT STEM. Ex: Zea mays.
T.S. of Monocot stem shows:
1) Epidermis: Outermost layer.Rectangular living cells are compactly arranged without intercellular spaces. On the outer surfacewaxy layer cuticle prevents evoporation of water from plant body.Numerous stomata in epidermis help in exchange of gases.Trichomes are absent.
2) Hypodermis: A clear cortex is absent. Formed of 3 to 4 rows of compactly arranged sclerenchyma cells,without intercellular spaces,giving mechanical strength to stem.
3) Ground tissue: Below Hypodermis,remaining part is extensively filled with soft parenchyma tissue called Ground tissue,with intercellular spaces.The tissue with chloroplasts synthesizes and stores food material.Endodermis is absent.
4) Vascular bundles: Numerous.Scattered in ground tissue and the arrangement is called Atactostele,an advanced character.Inner are bigger and outer are smaller in size.Each vascular bundle is called Fibrovascular bundle enclosed by a sheath of sclerenchyma fibres.Each vascular bundle is oval,conjoint,collateral,closed(cambium is absent ).Xylem is endarch(proto xylem towards center).Xylem shows two metaxyla,two protoxyla in Y shape.One protoxylem is crushed forming lysigenous cavity storing water.,forming protoxylem laculae.Xylem consists of tracheids,vessels,fibres,xylem parenchyma.Phloem consists of sieve tubes,companion cells.Phloem parenchyma is absent.Medulla ,medullary rays are absent.
Differences between internal structure of Dicot stem and Monocot stem.
Feature Dicot stem Monocot stem.
1) trichomes: present absent.
2) cortex: well developed.separated into highly reduced,
hypodermis,general cortex,endodermis. With only parenchyma.
Hypodermis: collenchymatous. Sclerenchymatous.
Endodermis: starch sheath. Absent.
3) stele:
Pericycle: sclerenchymatous. Absent.
Vascular wedge shaped,eustele. Oval,scattered,atactostele.
Bundles: Bundle sheath is absent. Fibrovascular ,
Sclerenchymatous .
Cambium: open closed.
Xylem: more,in serial order. 4, Y shaped,
2 meta xyla,2 proto xyla.
1 protoxylem into
protoxylem lacuna.
Lysigenous will not form. Stores water.
Cavity:
Phloem parenchyma: present. Absent.
--17—
INTERNAL STRUCTURE OF DICOT ROOT.
It shows:
1. Epidermis: Outermost living layer without cuticle and stomata.It is also called epiblema as it produces root hairs.Comparitively smaller epidermal cells forming root hairs are called trichoblasts.Epidermis gives protection and with root hairs help in absorption.
2. Cortex
igger than stele,showing:
a) Exodeermis: outermost layer, formed of 2-3 rows of thick walled suberised cells .It prevents exit of water from cortex.
b) General cortex: Below Exodermis, it is formed of several rows of thin walled living parenchyma.With leucoplasts it stores food. Thin walled cells help in conduction of water towards xylem.
c) Endodermis: Innermost layer.Single row of barrel shaped ,compactly arranged cells.Cells show deposition of lignin and suberin on the radial walls and transverse walls called casparian thickenings.Opposite to protoxylem,cells without casparian bands form passage cells,helping in the entry of water from cortex to stele.
3. Stele: Smaller than cortex, stele shows:
a) Pericycle: Single layered thin walled,rectangular cells formthe origin of lateral roots.Some cells form cambium during secondary growth.
b) Vascular bundles:Xylem and Phkoem alternate,on separate radii.Xylem exarch,with protoxylem towards pericycle.Xylem may be monarch(Trapa natans),Diarch (Nicotiana tabaccum),Triarch(Pisum sativum),Pentarch (Ricinus communis),Octarch (Castanea).Parenchymatous ground tissuein between xylem and phloem is called conjuctive tissue,helping in the storage of food material.
c) Medulla ; Small or absent due to union of meta xyla.If present,Parenchymatous storing food and water.
Differences between Dicot root and Monocot root.
Feature Dicot root. Monocot root.
1. cortex: relatively smaller. Bigger.
2. Pericycle: gives rise lateral roots, produces only lateral roots.
Vascular cambium,
During secondary growth.
3. Xylem: monarch to tetrarch. Polyarch.
4. Medulla: small or absent. Very big.
5. Secondary growth: occurs. Absent.
Differences between Stem and Root.
feature stem root
1. vascular bundles: collateral, conjoint. Radial.
2. endodermis: not clear very clear
3. xylem: endarch exarch
4.conjunctive tissue. Absent present
--18—
Transverse section of Dorsiventral ( Dicot ) leaf .
It shows three main parts:
I. Epidermis: Leaf is flattened,showing upper epidermis on adaxial surface and lower epidermis on abaxial surface.Epideermis is made up of single layer of barrel shaped cells ,compactly arranged without intercellular spaces.Externally covered by waxy layer called cuticle.More stomata on lower surface.Multicellular hairs are present on both surfaces.Epidermis protects inner tissues,helps in gaseous exchange by stomata.
II. Mesophyll: Tissue between upper and lower epidermal layers.It is hin walled parenchyma with chloroplasts.Seperated into upper palisade parenchyma and lower spongy parenchyma.
a) Palisade parenchyma: Below upper epidermis,1-3 vertical rows of elongated ,columnar cells,with narrow intercellular spaces.Larger number of chloroplasts are found nearer to cell wall.Helps in carbohydrate synthesis in light.
b) Spongy parenchyma: 3-5 rows of irregularly shaped parenchyma with large intercellular spaces,present towards lower epidermis.Intercellular spaces nearer to stomata are very large forming air cavities.Number of chloroplasts is less ,than palisade cells.Upper surface is dark green and lower surface is light green in colour.(dorsiventral leaf).Spongy parenchyma primarly helps in gaseous exchange,besidessynthesis of food to a lesser extent.
c) Vascular bundles: They are in mesophyll in the form of veins.Veins help in supplying water,salts, food,to allover the leaf surface,besides giving mechanical strength to leaf.Bigger vascular bundles at the base of leaf,smaller towards margins,apex.Vascular bundles are conjoint,collateral,closed.Xylem on upper side,phloem on lower side.Around each vascular bundle ,bundle sheath(border parenchyma) with compact specialized mesophyll cells form.Extending from the bundle sheath cells towards upper and lower epidermal layers,present bundle sheath extensionshelping in the conduction of food from mesophyll to vascular bundles.
TRANSVERSE SECTION OF ISOBILATERAL (MONOCOT) LEAF.
T.S.of isobilateral leaf shows:
I. Epidermis: On both upper side (adaxial side ) and lower side (abaxial side ) epidermal cells are barrel shaped.,compactly arranged.Epidermal hairs are absent.Waxy layer cuticle is present on the outer surface.Stomata are equal on both the surfaces.In grasses some enlarged,thin walled specialized cells filled with water are called bulliform cells or motor cells.These cells help in rolling and unrolling of leaf.Epidermis helps in transpiration and exchange of gases,besides giving protection to the inner tissues.
II. Mesophyll: The ground tossue present between the two epidermal layers is called mesophyll.It consists of columnar or spongy cells.,loosely arranged ,with intercellular spaces.They contain chloroplasts.Patches of sclerenchyma are
--19--
present beneath the epidermis,giving mechanical strength to leaf.Mesophyll is photosynthetic tissue.
III. Vascular bundles: Parrallel veins represent vascular bundles.They are conjoint,collateral,closed.Xylem towards upper eoidermis,phloem is below the xylem.Around each vascular bundle,specialized mesophyll cells,formbundle sheath.It may be rwo layered,outer chlorenchyma,and inner thick walled layerwithout chloroplasts.Schlerenchyma bundle sheath extensions ,on either side of vascular bundles,towards epidermal layers give mechanical strength to leaf.Bundle sheath cells with casparian thickenings on walls,are homologous to endodermis.
Differences between Dicot leaf and Monocot leaf.
Dicot leaf Monocot leaf.
1. stomata. More on lower epidermis. Equally distributed on both sides.
2. bulliform cells. Absent. Present on upper epidermis.
3. mesophyll. Differenciated into upper palisade undifferenciated.
And lower spongy parenchyma.
4. bundle sheath parenchymatous. Sclerenchymatous.
Extensions.